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In the Year F.C. (Future Century) 60, much of mankind inhabits space colonies which orbit the Earth. Dominance over the colonies is decided once every four years by a large tournament in which each nation sends a single representative to fight the others with a giant robot called a Gundam. Domon Kashuu is selected to represent Neo-Japan in one of these tournaments, but he fights less to ensure his nation's victory than to find his brother, who has been blamed for the deaths of Domon's parents and the disappearance of a very dangerous weapon, the Devil Gundam (Dark Gundam).

Six long months have passed. The Raalgon Empire has developed a horrible new type of weapon, and Tylor has been charged with the duty of intercepting it as it is being transported. But when all that could go wrong does go wrong, the crew members of the Soyokaze find themselves at the mercy of their enemies. As the hours tick down toward their execution, the crew wonders: has their irresponsible captain misled them? Or is this all a part of some greater strategy?

In 2099, Earth has shifted off its axis and begins to rotate at an angle of 90% to the 20th-century equator. Three years of natural disasters follows, and when things settle down, Earth's ten billion population has died and only two million moon colonists remain. Some return to begin the repopulation of the world in seven new cities and the remaining colonists of the moon fear that their former neighbors might pose a threat and construct a ring of defensive satellites to trap them on the newly repopulated homeworld.

Chirico is awoken from cold sleep and separated from Fyana. His attempts to find her come to the attention of a Nextant, the replacement for the Perfect Soldier program. At the same time, the new Pope is to be nominated, and Chirico is religiously considered to be "The Untouchable". With one of the nominees related to the Nextant, Chirico's actions will have far-reaching political ramifications.